The Outline of Science, Volume 1

When an animal frequents two different haunts, in one of which it breeds, it is very often safe to say that the breeding-place represents the original home. The flounder is quite comfortable far up the rivers, but it has to go to the shore-waters to spawn, and there is no doubt that the flounder is a marine fish which has recently learned to colonise the fresh waters. Its relatives, like plaice and sole, are strictly marine. But it is impossible to make a dogma of the rule that the breeding-place corresponds to the original home. Thus some kinds of bass, which belong to the marine family of sea-perches, live in the sea or in estuaries, while two have become permanent residents in fresh water. Or, again, the members of the herring family are very distinctively marine, but the shad, which belong to this family, spawn in rivers and may spend their lives there. So there are two different ways of interpreting the life-history of the salmon. Some authorities regard the salmon as a marine fish which is establishing itself in fresh water. But others read the story the other way and regard the salmon as a member of a freshwater race, that has taken to the sea for feeding purposes. In regard to trout, we know that the ranks of those in rivers and lakes are continually being reinforced by migrants from the sea, and that some trout go down to the sea while others remain in the freshwater. We know also in regard to a related fish, the char, that while the great majority of kinds are now permanent residents in cold and deep, isolated northern lakes, there are Arctic forms which live in the sea but enter the rivers to spawn. These facts favour the view that the salmon was originally a marine fish. But there are arguments on both sides, and, for our present purpose, the important fact is that the salmon is conquering two haunts. Its evolution is going on.[Pg 199]